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The National Aeronautical Collection originated in 1876, when the Chinese Imperial Commission presented the Smithsonian Institution with a gift of kites at the close of the Philadelphia Centennial Exposition. The collection expanded significantly after World War I and World War II, particularly because of the efforts of Paul Garber, who acquired some of the most important aircraft in the collection. These include the world's first powered aircraft, the 1903 Wright Flyer, Charles Lindbergh's historic transatlantic airplane, "The Spirit of St. Louis", and the first supersonic aircraft, the Bell X-1 "Glamorous Glennis". During these years the Smithsonian Institution and later the National Air Museum/National Air and Space Museum also collected numerous objects related to the history and technology of flight.

The collections of the Aeronautics Division currently comprise a total of approximately 28,000 objects, including 356 aircraft.

The major aircraft collections are: Early Flight; Commercial Aviation (includes piston-engine aircraft, turboprop aircraft, and jet aircraft); Military Aviation (includes pre-1920, interwar period, World War II, Cold War, and experimental); General Aviation (includes private aircraft, business aircraft, utility aircraft, gliders, kitbuilt and homebuilt aircraft, ultralights and hang gliders); Special Purpose Aircraft (includes research and experimental, exploration, record-setting, and racing aircraft); Vertical Flight (includes helicopters, autogiros, and vertical takeoff and landing aircraft); Lighter-than-Air Flight (includes balloons and airships).

The extensive non-aircraft collections of the division include: Aeropropulsion (engines, propellers, and auxiliary equipment); Armament and Ordnance (guns, cannons, bombs, bombsights), Art (includes space art); Awards and Memorabilia (trophies, medals and ribbons, insignia, memorabilia); Flight Material and Equipment (uniforms, personal equipment, including helmets, flight suits, etc.); Instruments, Avionics and Flight Management Systems (flight instruments, cockpit controls, and related devices); Kites; Model Aircraft and Engines; Photographic Equipment (aerial cameras); Posters.

The following online resources were compiled by Aeronautics Division staff: