This is the RL10, the world's first operational liquid-hydrogen/liquid oxygen high energy rocket engine and was re-startable in space. It was developed and built by the Pratt & Whitney Aircxraft Company. Two RL10s, each of 15,000 pounds of thrust, made up the Centaur upper stage of Atlas and Titan launch vehicles. A cluster of six RL10s also powered the second stage of the Saturn 1, a precursor to the Saturn V manned Project Apollo lunar launch vehicle.
The Atlas-Centaur first successfully flew in 1966 and was the first full-thrust re-start in space. Atlas-Centaur missions included Surveyor lunar probes, Mariners (Mars, Venus, and Mercury probes), and Pioneers 10-11 to Jupiter and Saturn. This engine was transferred to the Smithsonian in 1974 from the NASA Marshall Space Flight Center.
This object is not on display at the National Air and Space Museum. It is either on loan or in storage.